Category Archives: Uncategorized

Cranberry Lake Park: Golden Meadows Host Migrators…and Last Minute Mating!

The golden Eastern Meadow at Cranberry Lake Park in early September

Goldenrod! A variety of different Goldenrods gild Cranberry Lake Park in early autumn. Their bounty of nectar and pollen and the insects they attract make Cranberry Lake an ideal stopover for migrating birds and butterflies as they prepare for their long journeys. So cheer for Goldenrods as they feed wildlife, but don’t blame them for your fall allergies – blame ragweed!

Text and photos
by Cam Mannino

Year ’round birds and summer residents have almost finished molting and are also stocking up energy for winter or the migration. And a surprising number of insects are also preparing by busily mating one last time, leaving behind eggs or chrysalises that can bring forth young next spring and summer. Lots going on within this carpet of gold!

Migrating Birds –  Some Stayed All Summer; Some Just Stop Briefly for Rest & Refueling

Now’s the time to train your binoculars on any shaking leaf you see. It might be a fall migrator! Colorful, tiny warblers and vireos that are just passing through on their way south are well worth a pause to look into the shrubbery, as you’ll see below! But don’t forget to wish “bon voyage” to the migrators who arrived last spring and sojourned with us all summer. Many are finishing up their molts and readying for long journeys to the southern US or even Central and South America. We saw all of the birds shown below on the bird walk last week at Cranberry Lake Park.

Migrators from Farther North:  Just Passing Through

The trees are aquiver with fall migrators, travelers that bred up north and only briefly stop to rest and refuel around our woods and meadows. The most colorful ones that we often hear but can’t see are warblers and vireos. These little birds travel on the night wind because it’s safer. Hawks, eagles, and other birds that might see these small birds as food migrate during the day so that they can ride the thermals!

My camera and I are not quite quick enough to catch a lot of these tiny, fast-moving birds. So to share them with you, I’m relying on photos of other photographers with the skills, equipment and sometimes sheer luck to capture these little beauties!

The Blue-winged Warbler (Vermivora cyanoptera) breeds at Cranberry Lake Park, but the birds we’ve seen in the last week were probably moving through from further north. This fine bird shows off its black eyeline and blue wings with white wingbars in a gorgeous photo taken by gifted local photographer Joan Bonin:

Blue-winged Warbler (Vermivora cyanoptera) by Joan Bonin

This photo of the striking Black-throated Blue Warbler (Setophaga caerulescens) is by gifted iNaturalist photographer Jeff Skrent at iNaturalist.org.

A Black-throated Blue Warbler, photo by Jeff Skrent (CC BY-NC)

Yellow-throated Vireo  ( Vireo flavifronsfrom another generous iNaturalist photographer who uses the single name, paloma.

A Yellow-throated Vireo by paloma (CC BY-NC)

Nashville Warbler (Leiothlypis ruficapilla) was also too quick for me.  But here’s one I did manage to catch as it passed through in the autumn of 2016 on its way to Central America.

The Nashville Warbler on its way to Central America

Summer Visitors: The Migrators that Come Here to Raise Their Young

Some avian migrators see our parks as a great place to raise their young, so they come in the spring and stay for the summer. Having finished breeding and molting, they are now preparing to leave for points south. The male Common Yellowthroat’s  (Geothlypis trichas) “Witchedy – witchedy” song accompanied me often during the summer months as he and his mate raised their young. But you may here his “chuck” call as he hops among the branches stocking up on insect protein before leaving for Florida.

The Common Yellowthroat stops singing its “witchedy-witchedy” song in the fall and prepares to migrate.

Both the male Common Yellowthroat and his mate did a complete molt in July or August and now have fresh feathers for the trip. She’s bit more secretive than the male as she feeds on beetles, ants, bees (!), dragonflies and  grasshoppers within the golden fields. Usually she’ll leave a week or two later than the male.

The female Common Yellowthroat will be around a bit longer than her mate.

Last spring the Wednesday morning birders heard a Black-throated Green Warbler (Setophaga virens) singing its mating song along the Hickory Lane at Cranberry Lake. And this week, the birders spotted one flitting high in the trees. Perhaps it nested here to raise its young, or it could be just passing through our parks in spring and fall. Here’s  a photo I took of a male during the spring migration this year.

The Black-throated Green Warbler may have nested here this summer or he may be just passing through both in the spring and fall.

High in a snag, a Cedar Waxwing (Bombycilla cedrorum) paused briefly to survey the golden eastern meadow one afternoon. Many waxwings go into lower Canada to breed, but I’ve seen successful nests in our parks, too, so this one may have been a summer resident. Some will move south for the winter, but many waxwings will stick around during the cold months.

his Cedar Waxwing may have spent the summer with us or could be migrating south from Canada.

House Wrens (Troglodytes aedon) become quite secretive after breeding. But this one popped out of the greenery for a minute. It appears to have completed the late summer molt and is prepared to start south between now and mid-October.

The House Wren has raised its young here, molted and will be leaving shortly for Florida and other southern states.

A curious juvenile Field Sparrow (Spizella pusilla) peeked at me from dense shrubbery along the trail, too.  The migration of Field Sparrows is not always predictable. Some migrate, some don’t, and some migrate one year but not the next, according to the Stokes’ Guide to Bird Behavior (Vol.2). If this one does migrate, it may join a large flock with Chipping Sparrows as well as other Field Sparrows.

If this Field Sparrow chooses to migrate this year, it will probably join a larger flock. Or it may just stay put!

On one of my Cranberry walks, I spotted some Wood Ducks  (Aix sponsa) far across a well-hidden wetland. They were males who’d chosen this secluded spot to begin their molt out of the summer “eclipse plumage” into their fancy choosing-a-mate plumage. This one looked a bit rough at the moment, but well on his way to his courting colors. As I explained in last week’s molting blog, males molt earlier in the summer to camouflage themselves as females and then molt again in the fall, a time when they compete for a mate to breed with in the spring.

A male Wood Duck almost finished with the molt from his “eclipse plumage” to his courting feathers.

I often hear, but rarely see,  the vireos at Cranberry Lake. But I did get to see a Warbling Vireo (Vireo gilvus) in the spring. A rare treat, since it loves to sing high in the treetops! Its plumage is modest but its song is impressive!  The birders saw one lingering in our park for a while before departing for points south.

Warbling Vireos are often heard high in the trees, but not as commonly seen.

Occasionally I get a quick look at a Red-eyed Vireo (Vireo olivaceus). But Bob Bonin, Joan’s husband and another gifted photographer, got a much better photo than I have so far! This week it too was hanging out with other migrators.

Red-eyed Vireo – photo by local photographer Bob Bonin

Butterfly Migrators

Birds, of course, are not the only creatures migrating through our parks this autumn. Monarch Butterflies (Danaus plexippus) fluttered everywhere at Cranberry Lake Park in the last few weeks.  This “super-generation” of  fragile creatures – the ones that will fly 3,000 miles to Mexico – were swooping and diving over the meadows at Cranberry Lake, feeding on the acres of Goldenrod. So glad that we provide these master migrators with such a feast!

A female Monarch using it proboscis like a straw to sip nectar from Canada Goldenrod.

The tattered Giant Swallowtail (Papilio cresphontes) below looks lightly worn.  Let’s hope that attests to its having mated here this summer after its migration to Cranberry Lake.  These huge butterflies seem to be expanding their range north. Possibly due to climate change, Michigan now often has frost-free Septembers. So if mating was successful, the caterpillars of this very large butterfly may survive inside their chrysalises and hatch in our parks next spring.

This Giant Swallowtail looks a bit tattered at the end of a summer at Cranberry Lake Park. Due to mating? Maybe.

Last Chance for Progeny!  Insects Still Mating in the Meadows

Among the Goldenrod, insects seek out mates in a last ditch effort to leave offspring for next summer. A pair of Goldenrod Soldier Beetles (Chauliognathus pennsylvanicus) rendezvoused on a Bull Thistle  (Cirsium vulgare) – a potentially risky place to mate since a bird might think they make a tasty contrast against the blossom!

Goldenrod Soldier beetles mating on a Bull Thistle

A pair of tiny Pearl Crescent butterflies (Phyciodes tharos) also decided that a warm September afternoon provided the ideal time for mating.

A pair of Pearl Crescent butterflies mating in eastern meadow at Cranberry Lake Park.

And two Ladybugs chased around a Goldenrod stem, one frenetically holding on to the other. I couldn’t really determine whether they were mating or fighting! Clearly, one was more interested in escape and one was in hot pursuit. They were moving too fast for a great shot or a definite identification. My guess is that they were the invasive Asian Ladybugs (Harmonia axyridis). Unfortunately they are more common these days than our native Nine-spotted Ladybug (Coccinella novemnotata) and come in highly variable colors and patterns.

Two ladybugs chased each other around a Goldenrod stem – mating or aggression?

This tiny Red-legged Grasshopper (Melanoplus femurrubrum) had probably spent the night near the stalk of the Common Milkweed (Asclepias syriaca), wisely hidden from predators among its withering leaves. It will molt multiple times before mating and leaving eggs in the soil to emerge next summer.

A Red-legged grasshopper nymph probably spent the night within this Common Milkweed plant.

The edge of a meadow, near a wetland, might be an ideal spot for a female Great Blue Skimmmer dragonfly (Libellula vibrans). She could be spreading her wings in hope of attracting a mate. If successful, the aquatic plants nearby could host her fertilized eggs. Or while waiting for a likely male, she might just have a great perch for hunting unsuspecting prey!

A female Great Blue Skimmer sunned herself on a cool morning, possibly trying to attract a mate – or just hunting for the next unwary insect!

Spider Art On a Misty Morning

Early fall mornings are an excellent time to appreciate the art of the spiders. Warm days followed by cool nights leave heavy dew on spider webs, and when the sun makes them visible, they are dazzling. How about this lovely creation of an Orb Weaver spider (family Araneidae) drooping with the weight of the dew but subtly reflecting the colors of the sun’s spectrum!

The sun reflecting on the dew in an Orb Weaver’s web.

Another intriguing web, though not as beautiful, is cleverly constructed.  The Funnel Web Spider (Circuria species) lays a sticky sheet of web across the grass, which would be difficult to see were it not for the dew. At the edge of the web, it weaves a tunnel where it lies in wait for unsuspecting prey.  Above the sheet, it weaves an irregular network of silk designed to knock flying insects into its sticky net below. You can see the funnel at the back of this web below.

The Funnel Web Spider’s trap for flying insects with a nifty funnel in which the spider can wait for its prey unseen.

Frogs Underfoot!

As I skirted the edge of wetlands one Sunday, it seemed that a frog sprang out of the grass at every step! Most of them were Northern Leopard Frogs (Rana pipiens), their emerald green backs covered with circular black dots outlined in pale gold. Their dark eyes encircled with gold add to their glamour. Pinkish dorsolateral ridges are another distinguishing field mark. They shine like cloisonné in the deep grass!

Leopard frogs big and small sprung out of the grass near wetlands.

Until last week, I’d never noticed Pickerel Frogs (Rana palustris) at Cranberry Lake. These smaller frogs have somewhat rectangular spots that line up along the frogs’ back in neat rows. They are smaller than Leopard frogs. They tend to be brown (though they are sometimes green too), so I’m fairly sure this is Pickerel frog below.

Pickerel Frogs have more rectangular spots in rows down their backs, rather than the more random round spots of Leopard Frogs. So I think this is Pickerel frog, but I’m not positive!

And Finally, the Native Plants that Make It All Possible!

The plants, though not as often noticed by hikers, provide sustenance for all these creatures as the base of the food web. Butterflies sip their nectar. Bees and wasps feed pollen to their young. Other insects munch on leaves or make winter homes in the stalks. Birds, reptiles, amphibians, and mammals eat all parts of plants, or the insects that live on them. So here’s a gallery of  just a few of the plants that, along with the plentiful goldenrods, have bloomed in sun and shade to sustain the beauty and life of Cranberry Lake Park in late summer. (Use pause button for captions.)

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Autumn Mornings:  Not To Be Missed!

Mist rising in a meadow beyond the trees.

On our September bird walk at Cranberry Lake Park, we arrived on a cool fall morning. The previous day had been unseasonably warm, but a north wind sailed in overnight. The cool air had created heavy dew, leaving silver droplets that set the spider webs shining. The morning sun on the moist leaves created the fine mist you see above, rising  from a meadow beyond the Hickory Lane. What a sight! – the makings of future clouds floating like silver smoke above the wildflowers. A wave of migratory warblers and other small birds had arrived on that night wind. The birders watched, binoculars up, as these travelers hopped busily in the branches, feeding on the plenty of insects attracted by fields full of blooming fall wildflowers. And all of this beauty is gratuitous – unearned, just gifted to us if we just put on our shoes and head out the door! Consider escaping from the busy-ness of life for just an hour this week. Let these beautiful fall days lure you to the parks and savor the gifts that nature so generously offers to all of us.

Photos of the Week: Prepping for Change – the Fall Molt

An unidentified feather found in an earlier autumn. Anyone have a guess as to which bird? Perhaps a Downy Woodpecker? Mourning Dove? Flicker?  I’m stumped.

Maybe you’ve noticed individual feathers showing up in a nearby pond or on your lawn. Or you’ve noticed that birds and birdsong both seem a bit scarce right now. Or the birds you do see look just a bit scruffy. Well, don’t worry too much. It’s likely that nothing’s gone wrong for your avian neighbors. It’s just that lots of birds molt in late summer.

Text and photos
by Cam Mannino

That slightly disheveled bird you saw is probably an adult replacing frayed and worn feathers before the rigors of winter or before its fall migration. Once breeding is over, and while there’s still plenty of food to nourish them, birds systematically shed some of their feathers, replacing them before shedding others. Feathers account for 4-12% of body weight so it takes a lot of food energy to replace them! The foraging male Eastern Towhee (Pipilo erythrophthalmus) below with its feathers a bit askew is probably close to completing his molt. He will need to build up his reserves before beginning his journey to southern Ohio or points farther south.

A male Eastern Towhee almost finished with his fall molt and his foraging to restore his energy reserves.

So how does molting work? It turns out that feathers are made of keratin like our hair or fingernails, so feathers can’t repair themselves. The rigors of the breeding season can take quite a toll on feathers. Think of it  – all that to-ing and fro-ing from the nest or tree cavity, the scrabbling that occurs around an adult bird’s head as the young compete to be fed, plunging into the grass to snatch bugs, and riding down stalks to gather seeds on a windy day.  The feathers of an adult bird take quite a beating!  (Click on photos to enlarge; hover cursor for captions.)

When I saw this male Indigo Bunting (Passerina cyanea) a week or so ago, I wondered if he was just beginning his complete molt into the dappled blue and brown plumage that he wears during the winter.

A male Indigo Bunting looking more than a bit disheveled may be heading into his complete fall molt into brown plumage dappled in blue.

The molting process begins with the bird shedding a few feathers. In their place,  “pin feathers” emerge quickly. These pin feathers are supplied with blood for growth and look like the rigid shaft of adult feathers. As they lengthen, the blood stays in the lower ends and the tips slowly encase the feathers in a waxy coating. So when you see birds preening during a molt, they are very likely pulling off that coating, allowing the adult feathers to unfurl.

The Great Egret (Ardea alba) below seems to have shed a few feathers, since some were stuck to the branches behind it and some lay below on the trunk of the snag. Perhaps it’s unfurling new feathers, or it could just be going after some persistent feather mites.

A Great Egret preening. Is it molting or just getting rid of some feather mites? I’m not sure.

I’ve read that pin feathers are more sensitive than adult feathers. I wonder if that’s why we often see birds preening so vigorously before migration. Maybe preening feels something like scratching your head when it itches!  According to Cornell’s comprehensive “Birds of North America” subscription website, Egrets molt “some to most” of their body feathers when they’re in “northern non-breeding grounds.” Since egrets nest in only a few areas in Michigan, most of Michigan is a non-breeding area suitable for molting. Maybe that’s why this Egret was preening almost continually, including stretching up its leg to scratch its cheek!

A Great Egret who could be scratching at an itchy pin feather.

Molting happens on different schedules for different bird species. Some have one molt per year, some molt into breeding colors in the spring and molt again in the fall, and some do partial molts,  head and back feathers first and tail and wing feathers later. With experience the age and sex of many birds can be identified using their plumage.

In early summer after breeding, Wood Ducks (Aix sponsa) have an extra molt, a partial one. The males molt into “eclipse plumage, ” a drab plumage that makes the males look more like the brown females, though males still retain the red eye, red upper beak and double white stripe which females don’t have. Evidently male ducks do this to get the protection of being much less conspicuous during summer months. Here’s a photo of a male Wood Duck in eclipse plumage by iNaturalist photographer Heather  Pickard.

A male Wood Duck in eclipse plumage, looking a lot like a female Wood Duck. Photo by Heather Pickard  (CC BY-NC)

Like many ducks, Wood Ducks choose mates in the fall, overwinter with them and then mate in the spring. Of course the males want to look their most dazzling before courting! So they change into their breeding colors from mid-summer to early autumn, instead of in the spring like many birds. It’s a complete molt, meaning every feather is replaced, which renders them flightless for about three weeks. The female’s appearance stays the same with her new feathers. But the male recreates his elegant breeding plumage, including the iridescent green crest sloping down the back of his neck. Below are two males in a wetland near Lost Lake Nature Park two weeks ago who look like they’re about halfway between eclipse plumage and their glamorous breeding colors.

Birds don’t fight change. They simply do what’s required as the light informs them that the days are getting shorter. They molt. So when I go look for my jacket or warmer socks on a cool autumn day, I’m just part of the same process, I guess. And since my adjustment seems much less arduous than that of our feathered friends, I’ll try not to grouse about it!

[Footnote:  Information on molting was gleaned from Stokes' Guides to Bird Behavior by Charles Stokes (Volumes 1-3), this link at Wikipedia, Cornell Ornithology Lab's  subscription website www.birdsna.org. and this link at Cornell Ornithology.]

PHOTOS OF THE WEEK: Amazing Migrators on the Move!

Monarch butterflies at Tawas Point State Park last weekend. Photo by Nancy Isken

Well, they’re off!  When the wind sails in from the north, it’s a signal to all kinds of creatures: “Time to go!” So they take wing singly and in large flocks, letting the flow of cool air support them, carrying them quickly onward as they beat wings of all sizes to make their way to warmer climes.

Text and photos
by Cam Mannino

And of course,  it’s not just birds. Above you can see our friend Nancy Isken’s photo of  Monarch Butterflies (Danaus plexippus) last week resting at Tawas Point State Park before crossing Saginaw Bay. They were beginning their long journey to Mexico where they will spend the winter. In the spring, these Monarchs will make the first leg of the journey back north, stopping in warm areas like Texas to produce a new crop of butterflies who continue heading north. These new generations will fly only a few hundred miles, stop and reproduce, completing their whole life cycle in only  5-7 weeks. So it takes several generations to complete the trip  back to Michigan each summer. Sarina Jepson of the Xerces Society, which is focused on invertebrate conservation, says in a fine National Geographic article, “…when fall rolls around again, a special ‘super generation’ of monarchs that can live up to eight months will make use of air currents to wing all the way back to Mexico—a seemingly impossible feat for such a delicate-looking insect.” Imagine that!  So the Monarch butterflies born here In Oakland Township each summer can potentially live for 8 months instead of 5-7 weeks and fly 3,000 miles instead of a few hundred. We are living among  real, live superheroes!

A female Green Darner on the Wet Prairie along the Paint Creek Trail

And Monarchs aren’t the only insects that migrate. Green Darner Dragonflies (Anax junius) (also tracked by the  Xerces Society) travel down to Texas and Mexico as well. For dragonflies, it also takes many generations of reproduction before their trip is completed, though their migration is less well understood. But again, like the Monarch generations that arrive here each summer, the dragonfly offspring seem to know how to find their way in the right direction. Citizen science and research is helping explore dragonfly migration. But for now, I love a good nature mystery, don’t you?

Painted Lady butterflies (Vanessa cardui) make multiple generation migrations all over the world. But they migrate erratically. Some years they migrate and some years they don’t. And the direction and route can vary widely. Some experts speculate that their migration routes may be affected by dramatic changes in weather and climate – another nature mystery yet to be solved.

A Painted Lady sipping nectar during migration

At this time of year, the night sky begins to fill with thousands, even millions, of birds riding the wind south in the darkness, navigating by the stars or the setting sun, or by sensing the earth’s magnetic field. Some may navigate by landmarks or simply remembering good feeding grounds. And amazingly, most juvenile migrating birds, like those insects hatching during migration,  somehow know how to find their way without any help from adults! Cornell’s BirdCast website is a great way to watch the flow of birds across the United States day by day throughout the fall.

Remember all those bright little warblers, unusual sparrows and other small birds that sailed up here on a south wind in May?  Well, most of them have now finished breeding farther north and are beginning to make their way back to more warmth and sunshine. They’ve molted out of their bright breeding feathers so they’re a little less colorful now and their routes vary a bit depending on available food and weather. But keep a look out for these little travelers starting now.  Here are a few from the autumn of previous years:

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If you’re thinking you’d like to see some bird migrations in BIG numbers, you might check out the Hawk Fest featuring hawks, eagles, falcons and owls at Lake Erie Metropark on September 15 and 16. Or if you love our Sandhill Cranes (Grus canadensis), consider a trip to the Audubon Society’s  Cranefest at Big Marsh Lake in Bellevue, Michigan (near Battle Creek) on October 13 and 14.

A large flock of migrating Tundra Swans called over Cranberry Lake Park. (Photo by Bon Bonin)

Of course, if you want to stay close to home, you’ll be warmly welcomed at our Oakland Township Wednesday morning bird walks.  The walks rotate through our township parks each month and in September and October, they start at 8 a.m.  The schedule is available year ’round if you click on the drop-down menu above  for “Stewardship Events.”  We’re a friendly group with some experienced birders who are glad to help beginners see their first migrators.  Bring your binoculars or borrow a set from Ben, our Natural Areas Stewardship Manager, who leads the walks.

So yes, summer is waning.  But I can’t help feeling celebratory as autumn air turns crisp and the skies fill with winged creatures.  I recommend looking upward this fall and  perhaps wishing  “Bon Voyage!”, to our migrators who provide such beauty and mystery as they find their paths through the air.

Watershed Ridge Park: A Knee-Deep Immersion in Nature

The knee-deep flowers and grasses of a meadow at Watershed Ridge

Watershed Ridge Park offers an adventurous, challenging hike on a hot summer day. No trails exist yet to lead you through Watershed, since it’s not yet developed. But you can park by the evergreen trees just east of the maintenance shed and explore from there (approximately 1664 W. Buell Rd). This park is named for a watershed boundary, a high point within the park, that causes precipitation to flow in two directions, ultimately feeding both Paint Creek and Stony Creek. The diversity of habitats is remarkable. Large areas of the park have agricultural fields that are tilled and planted by a local farmer, which helps keep invasive plants out until the area can be restored to natural habitat.

Text and photos by Cam Mannino

But once I step out of the farm fields, the magic begins! A forest with vernal pools, a bright meadow, a boggy swamp shaded by trees, a seasonal stream, and a hidden marsh surrounded by a nearly impenetrable circle of native shrubs. So feel free to join me as I walk through some of this  wild and wonderful park on a hot summer day.

The Forest and Its Wetlands

I started down the edge of the farmer’s field to the west of the maintenance shed, walking carefully so as not to damage the Fogler’s soybeans. The field was dotted with wildflowers, like Pale-leaved Sunflowers (Helianthus strumosus) which attracted the attention of the Red-spotted Purple (Limenitis arthemis),  a butterfly that seems to be plentiful this year!

Pale-leaved Sunflowers shine in the shade under the trees that line the farmer’s field.
A Red-spotted Purple butterfly rests in the cool shade near the sunflowers.

Once I entered the cooler woods on the north edge of the field, I began to see blue-green wetlands shining in the dim, dappled light.

Blue-green wetlands glow in the distance as you enter the forest.

Oh yes, mosquitoes buzzed, but I patted on more insect repellent and headed for the water, because I knew that’s where I’d see the most wildlife. And sure enough, as I settled on a log near the water, a rustle behind me turned out to be a young White-tailed deer buck (Odocoileus virginianus) peering curiously at me from behind the greenery.

A young buck stares intently at me from the greenery near a wooded wetland in the forest.

I could hear an Eastern Wood-Pewee singing plaintively in the distance, a nice soundtrack for my entrance into a mysterious forest. A plop! at my feet made me aware of a young Green Frog (Rana clamitans), covered in Duckweed (genus Lemno), who’d been basking on a log just moments before.

A young Green Frog cools down among the Duckweed in a shady wetland.

Far out in the water, two Twelve-spotted Skimmer dragonflies (Libellula pulchellachased each other around a fallen willow until one finally settled for a moment. It was too far away for a good shot, so here’s my photo of a male in Bear Creek marsh several years ago.

A male Twelve-spotted Skimmer settles on a stalk in a marsh.

The forest wetlands were a busy location for courting that afternoon. A pair of Emerald Spreadwing damselflies (Lestes elatus) had evidently mated just before I arrived. Below you can see the female grasping the stem, her abdomen curled inward, preparing to lay eggs on a plant, while the male above keeps a wary eye for predators or other males.

The female Emerald Spreadwing Damselfly curls her abdomen to lay eggs on a plant while the male guards her from above.

It was hard to see how glamorous these Emerald Spreadwings are in the dim light, so I was pleased when a male landed in the sunlight nearby.

A male Emerald Spreadwing stops in the sunlight for a moment.

One of the stumps in the wetland was decorated with the wheel-shaped web of an Orb Weaver Spider (family Araneidae). It glowed in a ray of sunlight. According to Wikipedia, “Many orb-weavers build a new web each day. Most orb-weavers tend to be active during the evening hours; they hide for most of the day. Generally, towards evening, the spider will consume the old web, rest for approximately an hour, then spin a new web in the same general location.” Very tidy, eh? That might explain why I never saw the spider and there was virtually no detritus in the web. I’m not sure what to think about the extensive web on the stump below. A practice run?

The wheel-like web of an Orb Weaver spider

At the water’s edge, three “conks” of  Shelf/Polypore fungi shone white against the tracks in the wood, maybe the feeding galleries of emerald ash borer larvae .

Three shelf fungi “conks) on a log traced by a tunneling bark beetle.

Near another shady vernal pool wetland, I spotted a tiny Wood Frog (Rana sylvatica) gazing toward the water from a small piece of wood. Perhaps it was a fond last look since this little creature will soon move uphill a bit from the forest pool in which it hatched. Finding a place to hibernate under the leaf litter, it will freeze solid until spring. No heartbeat, no breathing, no movement. This seeming miracle is made possible by a glucose anti-freeze of sorts produced by its liver. As regular readers of the blog know, this strategy for survival never fails to astonish me.

Perhaps  this  tiny Wood Frog is contemplating its winter hibernation when it will freeze solid.

As I neared the edge of the woods, where it opens to the meadow, familiar plants were there to greet me.  I could have wished for less attention from Enchanter’s Nightshade, (Circaea canadensis) which at this time of the year is producing seed within burr-like fruits. Looking high into the trees, I strayed into a large patch and my socks were covered. For the rest of my walk, I became a transportation vehicle for Enchanter’s Nightshade fruits!

Enchanter’s Nightshade lies in wait for passersby to carry its seeds away to new locations.  My socks, for example, make a fine carrying device.

Another forest edge plant, Jumpseed (Persicaria virginiana), spreads its seed in a different way. Once the seed is mature, any disturbance can send its seeds flying up to three yards!

Once Jumpseed (pink flowers) produces mature seeds, bumping into the plants will propel the seeds up to 3 yards away.

Another native wildflower that hugs the forest edge, Jewelweed (Impatiens capensis), is called Spotted Touch-Me-Not for the same reason. Its mature seeds also spring forth from the fruits when they are disturbed.

Jewelweed also throws out its seed when touched, earning its other name, Spotted Touch-me-not.

The Meadow that Slopes Down to the Marsh

The meadow that slopes down to a marsh at Watershed Ridge

Walking out into the hot, moist meadow, I found myself knee deep in dense grasses and flowers. The sunlit field was slashed with long dark shadows from the forest as I waded through the dense growth underfoot.  All around me, dragonflies patrolled the flower tops, since the presence of a meadow in which to show off their colors and flying skills and a nearby marsh full of vegetation is their perfect habitat for mating and laying eggs. Widow Skimmers and Meadowhawks (genus Sympetrum) were everywhere. [I suspect the Meadowhawks were Ruby Meadowhawks (Sympetrum rubicundulum) but I’ve learned they can be confused with others unless you capture them and look closely – which I’m not at all interested in doing!)

A female Widow Skimmer displays against a grass stem.
A female Meadowhawk in bright sunlight cools herself by positioning her wings and abdomen.
A male Meadowhawk nearer the marsh spreads his wings to attract a mate.

Tiny butterflies fluttered through the flower stems adding their bit of color to scene. (Click on photos to enlarge; hover cursor for captions.)

As I struggled down toward the marsh, lifting my knees high to get through the lush tangle of stems, I begin to notice the plants near the marsh that love having “wet feet.” Mine, actually, were getting a bit wet, too!

Hearing sharp chipping calls behind me, I turned around to find an adult Song Sparrow  (Melospiza melodia) warning its offspring to stay out of sight. This may be a female since they often begin their molt later than males do. The youngster stayed out of sight within a nearby bush.

The adult Song Sparrow warned its youngster to stay hidden with a chipping call.

Adult birds can be a bit scarce in late July and August because many are molting a whole new set of feathers. They’re generally not a pretty sight in the process, I can tell you! They’re not hiding out of vanity, of course; they’re just more vulnerable as wing feathers are replaced. I could hear the “witchety, witchety” call of the Common Yellowthroat (Geothlypis trichas)beyond the wall of native Gray Dogwood (Cornus foemina) which prevented me from seeing into the marsh. And a Gray Catbird (Dumetella carolinensis) “miaowed” from the Dogwood as well, making only a brief appearance. So I took the bird photos below in other parks at other times.

Ben’s Balancing Act in a Shady Swamp

Swamps are generally defined by ecologists as a forested wetland. Watershed Ridge Park has a beauty.  Ben VanderWeide, our township’s Natural Areas Stewardship Manager, put on his waders one afternoon and went to explore this habitat at Watershed Ridge while working on a plant inventory of the park. What an adventure! The water may look shallow, Ben told me later, but the muck beneath it can be very, very deep! To explore the swamp, Ben tested each step and only went into the water once he found firm footing. Otherwise, he stepped or jumped from one hummock to another. Some of the hummocks were made by  “windthrows,”  fallen trees uprooted by the wind which had become covered by vegetation. Others were made by stumps of trees that had probably been felled years ago and were now overgrown with plant material.

While moving from hummock to hummock, Ben spotted some interesting and unusual native plants. How about Duck Potato (Sagittaria latifolia), also known as Common Arrowhead? I know I’d never heard of that one before! Ducks and other creatures love to eat its submerged tubers which store nutrients for the flower’s regrowth and/or reproduction.  And since, we humans love our favorite tuber – potatoes –  why not the name Duck Potato?

Duck Potato, so named because ducks and others eat their submerged tubers.

Ben found another unusual little beauty in the swamp, a Wild Calla (Calla palustris). Wild Calla is the only species in the genus Calla which is within the family Araceae. That family includes what are commonly called Calla Lilies (though they aren’t true lilies or in the Calla genus) and our old friend Jack-in-the-pulpit. It appears that the Wild Calla in Ben’s photo below had already been pollinated because, like Jack-in-the-Pulpit, the flowers are disappearing, replaced by green, berry-like fruits that will turn red in the autumn. Such a find!

A Wild Calla whose flowers have already been fertilized .  The resulting green fruits will turn red in the autumn.

Ben also spotted a sedge that he’d never seen in our parks before. Sedges (plants in the family Cyperaceae) are a big family of plants that look like grasses or reeds, but have triangular-shaped stems instead of flat ones. Papyrus and Water Chestnuts are in the same family. What I enjoy about sedges is that they often have such interesting spikes, like the one Ben found, Tuckerman’s Sedge (Carex tuckermanii). It also appears to have finished flowering and started producing fruits.

Tuckerman’s Sedge, a grass-like plant in the Watershed swamp

Along with these rewarding finds in the Watershed swamp, Ben was treated to a snack of High-bush Blueberries (Vaccinium corymbosum) as he balanced precariously on top of those floating hummocks!

Ben was also rewarded with High-bush Blueberries as he explored the swamp.

His treat was only fair, really, since he had to carefully avoid the abundant Poison Sumac (Toxicodendron vernix) scattered through the swamp. While he came home with a small Poison Sumac rash, it was a small price to pay for discovering cool, new plants and wild blueberries as far as Ben was concerned. Poison Sumac takes the form of a shrub or small tree and  grows only  in very wet places like swamps and bogs. My thanks to Ben for exploring this Watershed habitat. I don’t think I’ll be pulling on my waders anytime soon…but I’m very glad he did!

Poison Sumac, photo by Mawkaroni at iNaturalist.org (CC BY-NC)

When sufficient rain falls, a small stream flows out of Watershed’s swamp and finds its way through the woods westward,  eventually feeding the marsh I spoke of earlier at the foot of the big meadow. It’s not an impressive stream, but it feeds the plants in the marsh, which brings the insects, which feed the birds…you get the idea. Nature depends on connections like that.

A small seasonal stream flows westward from the swamp to the marsh at the foot of the big meadow.

Time to Head Home

By now, my damp feet, my tired knees and my socks decorated with Enchanter’s Nightshade made me aware that it was time to leave.  So I trudged back up the meadow, through the dim woods where the Pee-wee still asked its question,  “Pee-weee?”, along the Fogler’s soy beans and across the lawn to the cornfield beyond the shed.  There a group of young Red-winged Blackbirds (Agelaius phoeniceus) hung out on the cornstalks, moving restlessly like a bunch of rowdy teenagers, while their weary elders probably went on molting while hidden in the bushes and fields nearby.

The cornfield became a gathering place for young Red-winged Blackbirds.

I was standing next to a large tree stump when a juvenile House Wren (Troglodytes aedon) popped out of the vines that covered it.  This young bird  was a bit older than the plush-looking little Wren I’d seen at Stony Creek Ravine a few weeks ago.  But it was clear that the Watershed wren had still not quite grown into its beak. Surprised at how close I was, it hopped nervously for a moment, looked away, looked back – and popped back into the leafy cover. Caution won out over curiosity.

My Watershed adventure complete, I headed home, content that I’d experienced the natural world in the same way I’d relished it as a child growing up on Lake George Road. Yes, my arm had a few itchy spots here and there, my tired knees felt wobbly and water squished in my shoes –   but I’d meandered on paths of my own making,  out of sight of other humans, a quiet part of something wild and much older than I – and yet close enough to the kitchen at home that I’d be there in time for dinner. Thanks for coming along.

Footnote: My sources for information, besides Oakland Township's Stewardship Manager Ben VanderWeide, are as follows: inaturalist.org; Allaboutbirds.org, the website of the Cornell Lab of Ornithology at Cornell University; Wikipedia; Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela; Butterflies of Michigan Field Guide by Jaret C. Daniels, A Guide to Bird Behavior Vols.1-3 by Donald W and Lillian Q.Stokes, and others as cited in the text.

 

Paint Creek Trail: Last Hurrah of Spring Wildflowers, Tiny Pollinators and Nesting Migrators

Since this week’s blog features a variety of wildflowers along parts of the Paint Creek Trail, I thought we’d start with a little visit to the creek itself, which is currently overflowing its banks. The Swamp Buttercup certainly seems pleased with all that moisture! (If you increase your volume, you can listen to the soothing sound of flowing water and birdsong!)

Well, with summer heat settling in, native spring flowers are producing a glorious finale to the season before making room for summer blooms. Spring wildflowers love the weaker spring sun sifting through bare branches. But hot sun and the increasing number of leaves mark the end of spring blooms. Soon these wildflowers will set their fruits, send it to the ground by means of wind, water or simple gravity, and their seed-based reproductive efforts will come to an end for the season.

Text and photos
by Cam Mannino

So before they’re all gone, here are the ones I saw last week on the Paint Creek Trail between Gallagher Road and Dutton Road. You may still be able to spy some of these native beauties as you walk or bike on a sunny afternoon for the next few days.

Discoveries on the Trail between Gallagher and Silver Bell Roads

Golden Alexanders make a sunny yellow blanket beneath the trees just south of the Wet Prairie between Gallagher and Silverbell.

Golden Alexanders (Zizia aurea – above) should still be glowing beneath the light shade of trees near the Wet Prairie after Memorial Day. These members of the carrot family are visited by many spring pollinators, but they can also also self-pollinate. You can see them near the parking lot on Gallagher Road and at other spots along the trail as well.

In the Wet Prairie, a more unusual little wildflower is happily sprouting where the sun is full and the ground is moist. Blue-eyed Grass (Sisyrinchium angustifolium) is really a wildflower, though its leaves look like grass. Best to look for these dainty flowers on a sunny day, since the flowers stay closed on cloudy days.

Blue-eyed Grass on the Wet Prairie is best seen on sunny days.

Another unusual little wildflower huddles under its much taller grass-like stem on the Wet Prairie. Star Grass (Hypoxis hirsuta) shines up out of prairie on a 6 inch stalk topped by a small (3/4 inch) blossom. The six stamens with their angular anthers add to its star-like appearance. Maybe the dew on the leaf in the photo will help you sense the size of this tiny jewel of a wildflower.

Star Grass in the Wet Meadow with dew on a nearby grass leaf

Near the creek, just north of the Wet Prairie, a burst of orange caught my eye and I discovered a native flower that was new to me, Golden Ragwort (Packera aurea).  This plant too has multiple methods of reproduction.  Each of the tiny florets that make up the protruding center of the flower will produce an achene, a winged fruit enveloping a single seed that will disperse into the wind. But ragwort also produces underground stems (rhizomes) aid its spread!

Golden Ragwort brightens the bank edge of Paint Creek near the Wet Prairie

A native, fire-adapted plant which was new to me has the unusual name Bastard Toadflax (Comandra umbellata) and this is not a flax or a toadflax plant.  A report on desertusa.com, claims it was named “toadflax” by the 16th century botanist, John Gerard, because the flowers “be yellow, having a mouth unto a frog’s mouth” and “bastard” because at the time meant “false,” indicating perhaps that it was not a real toadflax. This interesting prairie plant is “hemiparasitic” because it feeds through its roots, but also taps into roots of other plants to get nutrition and water using special structures called haustoria. Bastard toadflax reproduces by rhizomes and produces a small oily fruits enjoyed by birds and mice, who by eating them carry the seeds farther afield.

Bastard Toadflax puts out underground suckers that tap into the roots of other plants.

If you notice a splash of white far back in the Wet Prairie, it might be a patch of  Robin’s Plantain (Erigeron pulchellus), a tough, little native that thrives after prescribed burns. It’s a relative of Daisy Fleabane (Erigeron annuus), a common wildflower in our parks. Unlike Daisy Fleabane, though, Robin’s Plantain more commonly produces one bloom to a stem. Like the Golden Ragwort, it reproduces by both achenes and rhizomes.

Robin’s Plantain, a relative of the more common Daisy Fleabane. Both thrive after prescribed burns.

Early last week, the fluffy, white blossoms of native Choke Cherry (Prunus virginiana) hung  in long clusters along single stems called racemes. After warmer sun later in the week, they had begun to turn yellow. They don’t produce edible cherries, but their bitter fruit appeals to game birds, songbirds and woodpeckers, according to one of my favorite wildflower sites, www.illinoiswildflowers.info.

Choke Cherry trees/shrubs don’t produce edible cherries, but rather a bitter fruit that appeals to birds, but not humans. But their blossoms are beautiful!

All long the trail you’ll see Wild Strawberry (Fragaria virginiana) blooming. Now here’s one bloom you might love to see fading, because its ripening fruit is tiny, but delicious. Good luck getting them before the birds and beasts do, though!

When Wild Strawberry blooms fade, on comes the fruit. But you’ll be lucky to get it before the creatures do.

Discoveries on the Trail from Silver Bell Road to Dutton Road

Despite our late spring, many different native wildflowers were blooming along this section of the trail.  At the bottom of the stairs leading to the creek, just past the parking area, Swamp Marigolds (Ranunculus hispidus) shine bright despite the deep shade. As you saw in the video at the top of the blog, the wetlands along the stream are doing what they do best – soaking up the runoff after heavy rains. And Marsh Marigold is doing its part!

 Farther down the trail, the lavender spikes of Wild Lupine (Lupinus perennis) are blooming. This beloved native produces seedpods open explosively, launching seeds up to seven feet from the plant. Lupine also produces colorful clones as it spreads with rhizomes. Lupines are rising toward the sun in several parks as part of the restoration work being done on our prairies. It’s so heartening to see these lavender blossoms opening up from bottom to top on their stately racemes. And I love the whimsy of its wheel-shaped leaves.

Both the leaves and blossoms of Wild Lupine make it a treasured native wildflower.

Along the berms on the western side of the trail, a perky yellow flower blossoms in bunches. The fuzzy stems and leaves give it the first part of its name, Hoary Puccoon (Lithospermum canescens). According to Wikipedia, the second word “puccoon” comes from an ancient Native American word “poughkone,” and refers to plants formerly used for dyes by Native Americans. I wonder if its roots can turn things that vivid yellow/orange?

The roots of Hoary Puccoon were used by Native Americans for dye. But please don’t pick them to try this at home!

Near the first bridge going south, I found a whole group of native wildflowers that were just finishing their blooms. A small group of Common Trillium (Trillium grandiflorum) still stood in the shade just off the trail. These flamboyant white flowers start turning pink as they age, eventually leaving only their leaves behind to gather up more sun for next years flowers. While true pink- and red-flowering trillium species exist, our Common Trillium just turn pink as they age gracefully.

Nearby Starry False Solomon’s Seal  (Maianthemum stellatum) bore only a rather disheveled version of its star-shaped blossom. After the blossoms, the plant will produce green fruits with purple/black stripes that eventually turn red. Woodland birds and mice enjoy the fruits, thereby carrying the seeds within the berries to new locations.

Starry False Solomon’s Seal has graceful leaves as well as tiny star-shaped flowers.

Native Bees and Tiny Butterflies Busy Pollinating

This may be a Mining Bee, a native pollinator, which specializes in Wild Geraniums.

Many spring wildflowers depend heavily on flies and native bees (of which there are 450 Michigan species!) for their pollination, because many other butterflies and the honey bees finish migrating or emerge from hives only in warmer months. Above is what I think is a native Mining Bee (Andrenidae family) foraging for pollen and thereby pollinating one Wild Geranium (Geranium maculatum) after another! I’m not knowledgeable about bees, but I learned from Pollinators of Native Plants that a particular Mining Bee, Adrena distans, is a specialist pollinator of Wild Geranium, so that’s my guess. Please feel free to correct me if you know more about bees than I do!

A few tiny butterflies were doing their part in pollinating flowers as well. If you see a fluttering wink of blue among the grass stems, it’s probably a Spring Azure butterfly (Celastrina Ladon). These 1 inch pollinators are blue when flying but once they settle, they quickly fold their wings so only the dull gray side shows. It makes them hard for hungry birds – and interested photographers! – to spot. Azures tend to pollinate Golden Alexanders, False Solomon Seal as well as other wildflowers. (Photo on the left by iNaturalist. org photographer Dan Mullen.)

The tiny Pearl Crescent (Phyciodes tharos) quickly fluttered by me and landed on dry grass stems in the Wet Prairie. These little butterflies produce several broods of caterpillars throughout the summer. So watch for tiny (up to 1.25 in.) flashes of orange and black rising from the grass and scurrying across trails all summer long, busily sipping nectar and inadvertently transporting pollen for the flowers.

The tiny Pearl Crescent can be seen now at the Wet Prairie and all summer long in our parks.

The last little butterfly was in a perfect spot to find a mate.  According to http://www.butterfliesandmoths.org, “To seek females, males perch in openings, flats or depressions near woods.” And that’s right where this one was – in the open, on a dead plant stalk near the trees just south of the Wet Prairie. He’s not glamorous but he has a glamorous name, the Dreamy Duskywing (Erynnis icelus). (Duskywings can be hard to tell apart, so again, feel free to correct me!) [Edit:  A man who collects butterflies and has seen many Duskywings told me on Facebook that he believes this is a Sleepy Duskywing (Erynnis brizo)  which he says are more common in this area.} He’s picked a great spot since one of his favorite nectars is the nearby wild lupine. These duskywings like to lay eggs on aspens and cottonwoods, which surround the Wet Prairie.

Possibly a Sleepy Duskywing  waiting on an open perch to be found by a mate.

Migrators Building Nests and, What Else? – Singing!

Late spring is a busy time for birds.  Some have just arrived and are busy singing to attract mates. A Warbling Vireo (Vireo gilvus) sang over my head in one of the Quaking Aspens (Populus tremuloides) near the Wet Prairie. This vireo is rarely seen once the leaves come on the trees; they like to stay high in the treetops and search diligently for caterpillars. But hearing a quick, rolling song high above me, I spotted his vireo shape, gray/olive back and yellow-ish white belly from below. Then he disappeared into the tree canopy. So here’s a photo of one I saw at Cranberry Lake a week ago. Click here to listen to his rollicking warble!

A Warbling Vireo sang above my head near the Wet Prairie. An illusive bird high in the trees with a great song!

The Warbling Vireo near the trail seemed quite frantic, singing insistently and hopping higher and higher in the tree as he did so. At first I thought another male vireo was singing across the trail from him – and I believe that he thought so too! But when I followed the other song, it was one of the great bird mimics, the Gray Catbird (Dumetella carolinensis), doing an imitation of the Vireo’s song and, I think,  driving him crazy! The Catbird was, as usual, adding a few flourishes of his own, but it seemed as though the Vireo found his call an annoying form of competition! For a sample of the Catbird’s wonderfully complex, mimicking song, click on the word “Song” at this link.

The Catbird’s mimicry of the Warbling Vireo’s song seemed to fool the vireo into thinking he had competition!

A pair of Baltimore Orioles (Icterus galbulaswooped down onto the Wet Prairie one afternoon. The female immediately disappeared into tall grass. It’s likely that she was carefully choosing grass stems out of which she could strip fibers for weaving her long, sack of a nest. Evidently, the male’s job was to stand guard while she had her head down.

A male Baltimore Oriole stands guard while his mate searches for just the right plant fibers to weave her bag-like nest.

Suddenly, the female oriole lifted out of the grass, followed immediately by the male, and they swooped into a tall Cottonwood (Populus deltoides) on the northern side of the prairie. She disappeared into a clump of leaves hanging high over the meadow, but I could see the branches moving as she worked and a tiny bit of the growing nest  (look in the center of the photo below through the leaves). I found a great little YouTube video of the Oriole’s weaving process, which is really incredible. Imagine tying knots and twirling fibers around a branch using only your beak!

The beginnings of an oriole nest high over the Wet Meadow

I heard a pair of  Eastern Kingbirds (Tyrannus tyrannus) fluttering together from tree to tree and stopped to watch. After giving what sounded like greeting calls, they soared up into a tall tree near the Wet Meadow. The female stood at the edge of a very messy nest which she was constructing on top of what appeared to be an old squirrel nest. The male doesn’t help build the nest, so while she worked, he stood guard in a nearby tree. I read in the Stokes Guide to Bird Behavior (Vol. 1) that Kingbirds build preliminary nests before settling into a final one to lay eggs. I’m guessing this may be just a trial nest, since building on a squirrel nest seems like an iffy location! (Squirrels are known to eat bird eggs!) The fine fibers drooping down over the top of the squirrel’s dry, gray branches and leaves are the female kingbird’s doing.

A female Eastern Kingbird seems to be constructing a preliminary nest on top of an old squirrel nest.

Kingbirds are identifiable from a distance by their upright posture – what one member of the birding group calls their “military stance.” And note the white tips on the tail feathers.

The male Eastern Kingbird stands guard while the female works on her nest

Every Nature Walk has its Special Moments

Last week, I had to smile at this seemingly relaxed male Northern Cardinal (Cardinalis cardinalis). He hopped slowly up and down a slope by the trail, picking up and rejecting items he found between the grass stems. With the grass taller than his head and his red crest (which is usually erect) drooping backwards, he reminded me of an avian Red Riding Hood!

It’s little things like that – and seeing the Blue-eyed Grass staring up at me – or watching the two Eastern Kingbirds near their nest – that send me home happy from every foray into the natural world. And these grant adventures only require the right clothes for the weather, water-proof shoes, a little application of bug spray at times and my undivided attention for an hour or so. I usually take my camera along,  but walks without anything but my trusty binoculars are just as rewarding. If you ever feel a bit weary as the poet Wordsworth once did, “The world is too much with us, late and soon/Getting and spending, we lay waste our powers…,” try a foray into our parks with your curiosity set at high alert. I’m confident that nature will send you home refreshed and with moments to share with others. Thanks for sharing mine!

Footnote: My sources for information, besides Oakland Township's Stewardship Manager Ben VanderWeide, are as follows: Caterpillars of Eastern North America by David L. Wagner; inaturalist.org;Allaboutbirds.org, the website of the Cornell Ornithology Lab at Cornell University; Wikipedia; Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela;Butterflies of Michigan Field Guide by Jaret C. Daniels,  A Guide to Bird Behavior Vols.1-3 by Donald W and Lillian Q.Stokes, Pollinators of Native Plants by Heather Holm,and others as cited in the text.