Tag Archives: Ladies’ Tresses

Watershed Ridge Park: Restoration Off to a Colorful Start!

 

Wildflowers re-establish themselves in a meadow at Watershed Ridge after invasive shrubs are removed

Watershed Ridge, at the corner of Lake George and Buell Roads, is not yet an accessible park. Trails have yet to be created for exploring its forests, wetlands and meadows, though they are planned for the future. Large areas of the land are still under cultivation for agriculture; local farmer, John Fogler, rents fields within the park to plant soy beans and pumpkins. (Click on photos to enlarge; hover cursor for captions.)

But last fall, our Natural Areas Stewardship Manager, Ben VanderWeide, began removing an almost impenetrable cover of invasive bushes that had taken over part of this beautiful piece of land. And suddenly, the graceful contours of the land came into view.

The slopes of Watershed Ridge after last fall’s removal of invasive shrubs

The rolling slopes looked lifeless then. But what a transformation this spring and summer as sun finally reached the native wildflowers!

The sloping landscape of Watershed Ridge Park this summer after the removal of invasive shrubs

Part of the reason Ben chose this area for restoration was that he’d noticed unusual native plants struggling to survive in the shade of invasive bushes like autumn olive, glossy buckthorn, privet, and such. Last autumn, Smooth Aster (Symphyotrichum laeveand a small orchid called Ladies’ Tresses (Spiranthes species) thrust their stems into the cool air. (Here’s a photo of the little orchid as it looked at the Wet Prairie on the Paint Creek Trail.)

Ladies Tresses, a small orchid, is a fall wildflower that Ben saw at Watershed Ridge last autumn.

Right now I’d advise anyone to be cautious about  walking through Watershed Ridge since it’s easy to get disoriented on 170 acres without trails (though a compass in your phone helps, I find!). Hikers need to use the edges of the farm fields so that they don’t damage the crops. The woody debris on the ground makes walking a bit tricky  and there’s a fair amount of Poison-ivy (Toxicodendron radicans) to avoid. Photos below might help those afraid of all 3-three-leaved plants. The one on the left is poison ivy.  Note the “thumb” on the leaves which is missing from other three-leaved plants like the Black Raspberry (Rubus occidentalis) on the right. Poison-ivy often has reddish “feet” when it’s climbing a tree and grows along the trunk rather than around it.

So for now, perhaps consider an easier alternative.

Blog post and photos by Cam Mannino

Join me on a virtual walk from the entrance area with its busy young birds, into the forest with its woodland flowers and wetlands and beyond the woods to the meadow with a riot of native wildflowers, butterflies, an exotic moth and more!

 

 

 

Birds Grace the Simple Entrance off Buell Road

The best spot to park is on Buell Road at the firewood pickup site (1650 W. Buell Road is the approximate address – click here for a map). Just to the west you’ll see a work area for Parks and Recreation staff that includes a pole barn and bulk materials storage. This work area is closed to the public, but you’re welcome to explore the rest of the park.

The old fence rows near the entrance on Buell Road have many trees, shrubs and vines, so birds nested happily around the area this summer. In mid-July, this area was full of fledglings trying out their wings and pestering their parents. A flock of young Eastern Bluebirds (Sialia sialis) settled on limbs and fence posts, begging to be fed while their beleaguered adults either surrendered or tried to escape their noisy youngsters. (Click on photos to enlarge; hover cursor for captions.)

A young Eastern Phoebe (Sayornis phoebe), however, was off on its own, trying out its wings, exploring the pole barn (sheds are popular Phoebe nest sites) and flying down into the grass like any adult Phoebe might.

A young Phoebe exploring the shed at Watershed Ridge – a very typical behavior for a bird that often builds nests in human structures.
The same Phoebe in a nearby tree to provide a glimpse of how small this little flycatcher is!

A young Downy Woodpecker (Picoides pubescens) was also off on its own and looking a bit dubious about what to do next as it sat in a nearby pine. This may be a young male since it appears that its juvenile red cap is slowly receding into the red spot that a male sports on the back of its head.

A juvenile Downy Woodpecker considers what to do next.

I failed to get decent photos, but two weeks ago, a pair of Cedar Waxwings (Bombycilla cedrorum) were still feeding their nestlings, flying back and forth between a tree on the west side of the area to a large willow on the east where they probably were foraging for insects before returning to the nest. Here are two not-so-hot photos!

From mid-July to mid-August, American Goldfinches (Spinus tristis) raise their young. Unlike most native birds, they wait until mid-summer to breed, because they like to line their nests with thistle down, eat the seeds themselves and feed them to their young. Several were flitting among the thistles along Buell Road.

American Goldfinches like thistle down for their nests and the seed for feeding themselves and their young.

A male Northern Flicker with his dashing “mustache” perused the territory from the top of a snag.

A male Northern Flicker with his dashing “mustache”

A “High Quality” Woodland Sprinkled with Wildflowers that Love Moisture, Shade and Rich Humus.

When you step into the cool darkness of the woods on a warm summer afternoon, you notice that most of the plants bear modest little flowers compared to sunnier areas. Perhaps that’s because they’ve  evolved growth and reproductive strategies that are different from the large, colorful blooms of a sunny meadow or prairie.

One good example is the unusual flower of a parasitic plant (not a mushroom!) that Ben helped me find, called Indian Pipe (Monotropa uniflora). This small, white wildflower typically grows in forests and lives on the photosynthesis of trees – especially oaks in our area – by tapping into the  mycorrhizal fungi that connect the trees below the soil. As a result, it doesn’t use photosynthesis and contains no chlorophyll. According to the Illinois Wildflower website (a descriptive floral site), “This unusual wildflower is normally found in high quality woodlands” where there is “abundant humus.”  Hooray for Watershed Ridge, eh?

The modest blooms of summer woodland plants primarily attract small bees, like sweat or carpenter bees,  or flies,  rather than honeybees or bumblebees.  They also frequently spread by rhizomes (underground stems) and/or by making bur-like fruits that will stick to any passing animal, including you!

Wildflowers that enjoy dappled light find a home where the woods meets the now-open meadow.

Naturally, mushrooms also thrive in shady, moist places. Near the edge of the woods, I saw what I think are two kinds of highly toxic mushrooms from the genus Amanita. I believe they are either Amanita phalloides or Amanita bisporigera. In any case, Amanita mushrooms are also called “death caps” for good reason –  so admire their exotic strangeness, but leave them right where they are. The yellow one may have been nibbled by squirrels or rabbits who can eat the toxins with no problem.

Wetlands Glow Blue/Green in the Deep Shade of the Woods

One of the wetland pools within the woods at Watershed Ridge

Two wetland pools shine in the darkness on either side of a ridge within the woods at Watershed.  Near the western one, an old log is festooned with a huge number of tiny, stalked mushrooms which I can’t identify. They make quite a show against the dark, wet decaying log on which they are thriving.  (Anyone have an ID for me?)

Ben saw two snappers on a log in one of these woodland pools, but they dove in before I saw them. I did see some of my favorite frogs, though. A small female Green Frog (Rana clamitans) (left) basked on a fallen log, while a male nearby (right) did the same. In the female, the tympanum (circle behind the eye used for hearing) is about the size of her eye; the male’s tympanum is about twice the size of his eye and he has a yellow throat.

At the eastern pool, a tiny, black-masked Wood Frog (Rana sylvatica), perhaps 1/2 inch long, paused for its photo on a leaf. This one’s back and legs are bright green, perhaps from the Water Meal (Wolffia columbiana) or the algae in the pool nearby. It seemed an odd time of year  to find an early spring frog, but Wikipedia says that  occasionally Wood Frogs do breed more than once per year. Maybe the heavy rains contributed to more water and hence more breeding behavior? Just a guess.

A tiny Wood Frog pausing on an oak leaf near one of the woodland pools

At the edge of the soybean field just beyond the wood, every step scared up hundreds of small Northern Leopard Frogs (Rana pipiens). They sprang out of moist grass and huddled beneath the bean plants. These young frogs  take about 70-110 days to develop after the females lay their eggs in the water between May to June. At about 2 inches long, they’re half the size of a mature Leopard Frog. Their numbers have generally been in decline since the 1970’s so what a delight to see so many of them here in the township on a protected piece of land!

A small (about 2 inch) Leopard Frog, among hundreds that sprang out of the tall grass into the bean field a few weeks ago.

A Riot of Color as Wildflowers and Butterflies Reclaim a Wild Meadow

Black-eyed Susans and Butterfly Milkweed “take the field” after invasive shrubs are removed from Watershed Ridge

Step out of the woods and the colorful signs of restoration engulf you. Once shaded-out by invasive shrubs, native wildflowers like Black-eyed Susans (Rudbeckia fulgida) and Butterfly Milkweed (Asclepias tuberosa) in the photo above sprang back into the sunlight that finally reached them.  So exciting to see these plants come back with such vigor in the first season after the invasives are removed! (Use pause button to read captions.)

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And with the wildflowers, of course, come the butterflies – but first, one glorious moth! When I first spotted this unusual creature, I thought it must be some sort of bumblebee. But no, it’s the Snowberry Clearwing Moth (Hemaris diffinis). It has fewer central scales on its wings than other butterflies or moths so the wings appear clear in places. It’s odd shape and hovering flight have also inspired the names “the hummingbird moth” (not to be confused with the European one of the same name, Wikipedia tells me!) or the “flying lobster!”  A new creature for me!

This is not a bumblebee. It’s a Snowberry Clearwing Moth mimicking one!

On my first summer visit with Ben, we saw a glorious Eastern Tiger Swallowtail (Papilio glaucus) sipping from some of the bee balm which emerged in great waves of color across the restored meadow this summer. What a sight!

A female Eastern Tiger Swallowtail sips nectar from the great swath of bee balm that now flows across the restored meadow.

Having seen the Spicebush Swallowtail butterfly (Papilio troilus) for the first time at Charles Ilsley Park a few weeks ago, I saw another male at Watershed Ridge.  The clue is the wash of blue scaling on the wings. A week later I saw another one near the shed which was either the female, which has a less intense wash of blue scaling, or a tattered male who had already lost some of his scales. That Spicebush continually tried to chase off a female Monarch (Danaus plexippus) who was patrolling the same patch of flowers last weekend. The scales on her wings looked quite worn as well. Perhaps they’d worn each other out with all that chasing! I saw some fritillaries dancing together at a distance but not close enough for a shot. But the little Orange Sulphur  (Colias eurytheme) was kind enough to pose for a few moments.

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The meadow slopes down to a sunny wetland that is probably fed by the creek running through the trees just east of the meadow. The stream bottom is bright orange, most likely a result of iron deposits in the soil. I loved how the sky was caught in the water’s surface as I looked upstream.

The resulting wetland at the meadow’s edge featured flowers that love “wet feet,” like Joe Pye Weed (Eutrochium maculatum) and Boneset (Eupatorium perfoliatum).

With a wetland nearby, dragonflies, flying predators, swooped across the meadow on the hunt for other insects.

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A Meadow Under Restoration Gets Us Dreaming

Another view of the meadow that is slowly being restored at Watershed Ridge

Witnessing the dramatic renewal that takes place when invasive shrubs are removed from park land is simply inspirational. What else, who else might return to this lovely piece of land once the Parks and Recreation Commission has time and the funds to develop this 170 acres? What seeds are waiting in the seed bank for their turn in the sun – perhaps even this autumn when the asters in their varied lavender-to-purple blooms or other autumn wildflowers rise from the earth? What other exotic creatures like the Snowberry Clearwing Moth might be sipping at future flowers? What birds might return to nest here? The possibilities are endless as the Ben and the Parks Commission work to renew the diversity of life which is our township’s natural heritage. So much to look forward to as this park progresses over the next several years!

Footnote: My sources for information, besides Oakland Township's Stewardship Manager Ben VanderWeide, are as follows: butterfliesathome.com;Pollinators of Native Plants by Heather Holm; Stokes Nature Guides: A Guide to Bird Behavior Volumes 1-3; Allaboutbirds.org, the website of the Cornell Ornithology Lab at Cornell University; Wikipedia; Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela; other sites as cited in the text.

OUT AND ABOUT IN OAKLAND: Nature and Park Stewardship Working in Harmony All Over the Township

Blog and photos by Cam Mannino
Blog and photos by
Cam Mannino

From early autumn until the first really cold weather,  nature is busy preparing for winter – and so is the Parks Commission. While plants disperse seed and mushrooms release spores for next spring’s growth,  our stewardship manager, Ben VanderWeide, is sowing native seed as well. As the trees drop leaves to prepare for spring growth, Ben and his contractors clear away invasive shrubs to provide spring sunlight  for native  grasses, bushes and wildflowers. So this week, please join me for some short visits to several parks to see how nature and the Parks Commission work together to prepare for winter snow and the spring to come.

 

Bear Creek Nature Park: Seeding, Feeding and Choosing a Mate

Bear Creek made a glorious exit from autumn. In late October, it burned with autumn gold and red.

Fall at Bear Creek
Late October at Bear Creek Nature Park

As usual, the Black Walnuts (Juglans nigra) “fell asleep” before the other trees; they “wake up” late in the spring, too. I like to think they need more rest than other trees!

Walnuts sunset BC
Bare trees on the Walnut Lane at sunset in Bear Creek

The Mallards (Anas platyrhynchos) were busy with two kinds of preparation: finding a mate for the spring  and eating to store up fat for the winter. Happily, both could be accomplished at once. Males, now in courting colors, cruised the Center Pond with females, going “up tails all” while feeding below the surface.

Mallards Up Tails All BC
A small flock of upside down ducks foraging underwater at Bear Creek’s Center Pond.

Mallards choose partners in the fall and then mate in the spring. So eventually they begin to pair off like these two did as the setting sun gilded the pond.

Mallard Silhouette Sunset BC
A mated pair of Mallards float through the golden light of sunset on the Center Pond

Plants, of course, are dropping seeds and fruits. The white fruits of Gray Dogwood (Cornus foemina) have either been eaten by birds or dropped to the ground to produce a dense thicket of more dogwood next year. On the right are the bare red pedicels in November. (Click to enlarge; hover cursor for captions.)

At the edge of the wood, I spotted a Gilled Mushroom (order Agaricales) within a dead stump. It will drop its spores from the gills beneath the cap to produce more mushrooms next year.

Mushrooms piled in stump BC
Gilled mushrooms will drop spores from the gills beneath their caps

Lost Lake Nature Park:  Shedding Leaves and Seeds

Forest on LL 1
The forest behind the sledding hill in late autumn

Fall is a time when trees shed their summer leaves. The apparent path of the sun lowers toward the horizon from September to Winter Solstice (December 21). So,  the days grow shorter. Less daylight means that leaves lose the chlorophyll that makes them green and allows them to photosynthesize. This process of photosynthesis stores energy from sunlight in the chemical bonds of sugars. Trees use these sugars to grow during spring and summer, and many times sugars are stored for the future as starch. In the fall leaves change color as the chlorophyll recedes, their job completed for the season. Eventually lower light signals the trees to create “abscission cells” which grow between the stem and the branch, separating them.  Down come the leaves. (“Abscission,” by the way, has the same root as the word “scissors!”)

Leaf carpets are useful as well as beautiful.  As they decompose, leaves release nutrients back into the soil and provide the spongy humus that helps the soil hold water. And of course, they can act as protective mulch for the roots of forest plants and trees.   Here’s a colorful carpet of White Oak and Red Maple leaves near Lost Lake.

Bur Oak Leaf collage
A carpet of White Oak and Red Maple leaves near Lost Lake

Out in Lost Lake, the big flat leaves of Fragrant Water Lilies are fading for all the same reasons as other leaves. Their graceful simplicity against the dark water was eye-catching.

In the forest, near the top of the sledding hill, some native plants were still preparing to drop their seeds. Showy Goldenrod’s (Solidago speciosa) seeding plumes still stood tall in the forest light under the trees. And a stem of Bottlebrush Grass (Elymus hystrix) had yet to let go of  its awn-tipped seeds. The grass fruit flies through the air and lands on the ground like tiny arrows, carrying their cargo of next year’s seed.

A flock of restless American Tree Sparrows (Spizelloides arborea) swooped and dove near the western edge of the lake and foraged over the grassy area near the former stable. Tree Sparrows, summer residents of the Arctic, are experienced foragers. They spend the winter here finding edible seeds, fruits, insect larvae –  whatever they can find in a snowy Michigan winter.

Tree Sparrow LL
A Tree Sparrow makes a good winter meal from the seeds of sedges, grasses, insect larvae and whatever else it can find.

At the end of the walk, human prep for winter becomes apparent. Jeff Johnson, the Parks Commission’s Maintenance Tech, spent about a day and a half preparing the sledding hills for winter use. He marked off the big hill, the kiddie’s hill and the boundaries of the upward trek. Just in time for the big snowfall!

Sledding hill prep LL
The sledding hill was prepared for winter. And the snow arrived a few days later.

Cranberry Lake Park:  Pond Ice and Puffballs

In the center of Cranberry Lake Park is a large pond where I’d seen herons and mallards feeding together one early fall afternoon. When I went back to explore in late fall, the pond was beautifully still and silent.  A light skim of ice had formed on one end.

Pond CL south end w ice skim
A skim of ice formed on the pond at the center of Cranberry Lake Park

Near the pond, a large colony of tiny Puffball Mushrooms (genus Bovista) had finished releasing their spores. These tiny puffballs tend to grow in groups like this.

Puffballs on long b. Bov.ista
Tiny puffball mushrooms cover a log near the pond in the center of Cranberry Lake Park

A few years ago, I saw these little mushrooms covering a stump at Bear Creek.  They were plump in early autumn, filled with spores. When the spores are mature and are tapped by falling raindrops, small creatures or the occasional curious human – poof! – the spores pop out of the open center to be carried on the air so more mushrooms appear the following year.

Puffballs when the spores are developing are plump and sensuous.
Puffballs when the spores are developing are plump and sensuous.

The little mushrooms at Cranberry Lake  were spent; their center holes had opened, releasing the spores.  Quite a come-down in appearance, I’m afraid.

Once the center opens and releases the spores they look quite different!
Once the center opens and releases the spores,  these tiny Puffballs look quite different!

Stewardship: Lending a Helping Hand to Nature’s Winter Prep

The Wet Prarie on the Paint Creek Trail: Birds Gather Seeds and So Do We

The Tree Sparrow at Lost Lake is not the only creature foraging for wild seed this autumn. Ben and a crew of volunteers spent a peaceful morning at the Wet Prairie off the Paint Creek Trail gathering native grass and wildflower seeds for planting in other areas of the township.

Gallagher Creek and Charles Ilsley Parks: Seeds Sown by Plants and People

While the native plants are dropping seeds to prepare for spring abundance, so our Township Stewardship Manager, Ben VanderWeide is busy sowing native seed around the township. During the first week of December, parts of two parks, Gallagher Creek  and Charles Ilsley Parks,  were planted with native seed that Ben had gotten through a US Fish and Wildlife grant. The land had earlier been prepared in Gallagher Creek’s west section by clearing invasive shrubs and plants through selective treatment, mowing, and prescribed burns. Now a no-till native seed drill, designed to handle the varying sizes of native seeds,  went to work. Discs cut a thin furrow in the ground to a pre-determined depth and simultaneously, seeds were dropped into the thin furrows.

Seeding Gallagher Creek
Planting native seed at Gallagher Creek Park

At Charles Ilsey Park, the machine sowed more native seed after last year’s more extensive prairie planting.  Here’s a YouTube video of the native seed drill at work.

Watershed Ridge Park, the Wet Prairie and Stony Creek Ravine Nature Park:  Trees and Humans Prepare for New Growth

Just as trees prepare for spring by discarding their old leaves in the autumn, Ben and his contractors are working on ridding our parks of shrubs and underbrush to encourage new native growth. After cutting invasive woody shrub sprouts on the Wet Prairie, Ben carefully treated stumps to prevent them from re-sprouting next year. Invasives are persistent,  so Ben has to be too!

At Stony Creek Ravine Nature Park, Ben brought in a contractor with a forestry mower to clear invasive shrubs from  the current entrance. What a difference! Though not taken at exactly the same place, the photo just below will give you an idea of the density of the invasive shrubs at Stony Creek Ravine before removal – and below that is the wonderfully open look it has now.  A lovely view from the beautiful stone bench in the distance.

My husband in the dense invasive shrubbery at Stony Creek Ravine
My husband in the dense invasive shrubbery at Stony Creek Ravine Nature Park
The park entrance as it look after removing invasive shrubs
The park entrance as it looks after removing invasive shrubs. The Kezlarian stone bench is in the distance.

But the biggest, most impressive “clearing the decks” project this fall was the beginning of habitat restoration process at Watershed Ridge Park. Just a month ago, the rolling slopes of this park were tangled with invasive shrubs that in many places were impenetrable. However, Ben noticed sun-loving native grasses and wildflowers struggling to survive under the heavy cover of overhanging limbs and vines – plants like the delicate orchid Ladies’ Tresses (Spiranthes species), Large-leaved Aster (Eurybia macrophylla) and three kinds of native Goldenrod.

So the first week in December, restoration began. A contractor’s forestry mower cleared 10.5 acres of invasive shrubs from among the trees. Now the contours of the land can be seen, dotted with oak, ash, elm and cherry trees.

The land slopes down from east to west to a marsh and a pond, both fed by a stream running from the large marsh that’s to the north of the park.

Stream at Watershed Ben's photo
This un-named stream runs from a larger marsh on the north through the woods to a marsh and pond on park property.
Cleared Hillside Watershed Ridge
Looking south toward Buell Road across a newly cleared hill that slopes down to the pond.

Now the stream, the marsh and the pond, which extends as far as Lake George Road, can be approached easily without fighting through dense, invasive shrubbery and vines. Once trails are created, this will be a lovely spot to watch for water wildlife!

Pond at Watershed Ridge
The west end of the park’s pond that extends off  Lake George Road.

In the forest at the edges of the cleared area are Red, Black and White Oaks. Here’s an old White Oak (Quercus alba) within the cleared area. Probably because of the crowding from shrubs and small trees, it never had the chance to spread its limbs wide in the sunlight, like the White Oak we all enjoy near Bear Creek’s Center Pond.  But it’s still pretty impressive!

White Oak at Watershed Ridge
A large White Oak within the newly cleared area.

I couldn’t  resist taking a photo of the orange Turkey-tail Mushrooms (Trametes versicolor) Ben spotted sprouting from a fallen long nearby. Turkey-tails come in a wide variety of colors!

turkey-tail-mushrooms-watershed
Turkey-tail mushrooms on a fallen log

In case you’re interested, this is the contractor’s forestry mower that cleared the shrubs at Stony Creek Ravine and also at  Watershed Ridge.

Shrug-chewing machine WR
Forestry mower used at Stony Creek Ravine Nature Park and Watershed Ridge Park. Thanks to the Ruffed Grouse Society for your great work!

What an exciting beginning to restoring this piece of land and creating a park that the community can enjoy! It will take time to develop trails and any other amenities that are appropriate to this beautiful land with its woods, slopes and wetlands. We’ll try to keep you posted on developments.

Nature and Parks Stewardship: Partners in Fostering Our Natural Heritage

Stony Creek Ravine late autumn SCR
Stony Creek Ravine Park in late autumn

I’ve come to see that stewardship supports nature by mimicking it in so many ways. Seeds that create new growth each year drop to the ground or sail away on the wind to take root and grow into more life. And we gather native seed and carefully sow it in narrow furrows to do the same. Trees drop their leaves to make way for new ones next spring. We too clear away invasive plants and shrubs to make way for new life. Native seed, lying dormant in the underground seed bank for years, will now sense exposure to sunlight and moisture on the earth above, crack open and begin to sprout. And with those plants, eventually will come butterflies, other insects, and birds that eat or nest in those native plants, restoring the diverse habitat that is part of our area’s natural history.  So much life from simply giving nature a helping hand!

 

Footnote:  My sources for information,besides Oakland Township's Stewardship Manager Ben VanderWeide, are as follows: Ritland, D. B., & Brower, L. P. (1991); Stokes Nature Guides: A Guide to Bird Behavior Volumes 1-3, Allaboutbirds.org, the website of the Cornell Ornithology Lab at Cornell University; Wikipedia; http://www.butterfliesandmoths.org; Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela; Butterflies of Michigan Field Guide by Jaret C. Daniels; University of Wisconsin's Bug Lady at www4.uwm.edu/fieldstation/naturalhistory/bugoftheweek/ for insect info; http://www.migrationresearch.org/mbo/id/rbgr.html for migration info; invaluable wildflower identification from local expert, Maryann Whitman and the website www.illinoiswildflowers.info; experienced birder Ruth Glass, bird walk leader at Stoney Creek Metro Park for bird identification; Birds of North America Online; Audubon.org; Nature in Winter by Donald Stokes, Trees in My Forest by Bernd Heinrich, Winter World by Bernd Heinrich, Savannah River Ecology Lab (Univ of Georgia); Tortoise Trust website www.tortoisetrust.org;  An Orchard Invisible: A Natural History of Seeds by Jonathan Silvertown; The Ecology of Plants by Gurevitch, Scheiner and Fox; other sites as cited in the text.