Tag Archives: Eastern Wood-PeeWee

Charles Ilsley Park: Our First Restored Prairie in Bloom! And Oh, the Butterflies and Birds!

 

The Eastern Prairie at Charles Ilsley Park
Blog post and photos by Cam Mannino

Charles Ilsley Park sways with grass and colorful wildflowers right now as restoration begins returning the rolling fields to the prairies they once were. The eastern prairie, featured this week, is breath-taking!   Butterflies dance in and out of native Black-eyed Susans, bees hum, grasshoppers leap at your feet and dragonflies swoop above the Yarrow and Bee Balm. Birds perch and forage at the trail edges and sing from the woods.

These former farm fields, once full of invasive and non-native agricultural plants, are gradually being restored to provide for the birds, insects and other wildlife that historically nested and foraged here. A new path through the thirteen acres of the eastern prairie now provides a perfect quiet stroll on a sunny morning or afternoon.

Birds are Plentiful, Colorful and Sometimes Difficult to See

At this time of year, adult birds are busy feeding and over-seeing either nestlings or fledglings. So they’re deep in the leaves, high in the sky, or down in the grass, foraging for whatever will feed the young. Here’s a graceful female Baltimore Oriole (Icterus galbula) with seeds in her beak approaching her nearby nest hidden in a clump of leaves that hangs right over the trail as you enter the park.

A female Baltimore Oriole with seeds in her beak for her nearby nestlings

Earlier in the summer, the bird boxes along the trail into the park were busy places! A House Wren (Troglodytes aedon) took over a rather dilapidated one.  You’ll notice in one of these pictures, the wren is pecking at a spider web. Cornell Lab says that wrens often put spider egg sacs into their nest box so the spiders will rid it of mites! (Click on arrow for slide show; use pause button for captions.)

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The Tree Swallow (Tachycineta bicolor) spread its huge wings as it swooped in and out feeding its mate or young in June. Now the box seems to be just a stop-over spot as it soars and dives in the meadow beyond.

Look at the size of those wings!
And look at those wings folded!

Thanks to Laurie Peklo, a member of the birding group who cleaned out the nesting boxes this spring, the Eastern Bluebirds (Sialia sialis) also found a home at Ilsley again this year.

A male Eastern Bluebird guarding his nest box.

A few weeks later, we saw a Bluebird fledgling in a tree, perhaps the male’s offspring. Baby bluebirds take a while to turn really blue, so I’ve also included a photo of a bit older one begging in our front yard. (Click on photos to enlarge; use back arrow to return to blog.)

When you reach the sign about Prairie Restoration, you’ll see that the center of the park is brown and mowed. Don’t worry.  Ben VanderWeide, township stewardship manager, and his crew are just preparing the ground so that in the fall, it can be sown with native prairie plants – just as the eastern prairie was planted two years ago, and the north and west were planted last year. In a few years, we’ll have close to 50 acres of prairie in this park!

On every recent visit, a male Common Yellowthroat (Geothlypis trichas) has sung from the bushes on the hill in the center field. That’s why Ben left the greenery there when the field was prepared for prairie planting – to protect their young. The photo below, was taken in the spring when one male came a bit closer and really belted out his “witchedy, witchedy” song!

The “witchedy” song of the Common Yellowthroat can still be heard at Ilsley, but in spring he really threw back his head and SANG!

Field Sparrow (Spizella pusilla) foraged in the grass of the trail on one of our birding walks. Its pink-ish beak matches its pink legs – an easy field mark for these little birds. Ben describes their rolling song as a bouncing ball.

A Field Sparrow foraging at the edge of the trail

Young Chipping Sparrows (Spizella passerina) and their elders forage there as well. Thanks to birder extraordinaire, Ruth Glass,  for the identification! (Click on photos to enlarge.)

Ruth also identified the rather scruffy  Eastern Wood- Pewee fledgling (Contopus virens) below.  

A rather scruffy-looking, juvenile Wood-Pewee

We birders heard an adult one singing in the woods near the eastern prairie last week, but didn’t see it – as is often the case with singing Pewees.   Luckily, one landed near our deck at home a few weeks ago, singing its name on a rising note  – Pee-weeee! Nice to see one up close and personal!

An adult Eastern Wood- Pewee
Indigo Bunting

In a clump of trees on the south side of the east prairie, an Indigo Bunting (Passerina cyanea) sang from a very tall snag on my last three trips to Ilsley. He must have a nest nearby. He’s so far up, singing his song in phrases of twos and threes, that it’s hard to even see he’s blue!

But my husband and I were lucky enough to have one fly down to a small tree near us while walking at Addison Oaks about ten days ago. Such a glorious bird!

An Indigo Bunting – one of the rare moments when I’ve seen one up close!

As some of the birders exited the eastern prairie last Wednesday, they saw a very unusual bird, a Yellow-throated Vireo (Vireo flavifrons). I didn’t get a look before it flew – so here’s a lovely photo from a generous Creative Commons photographer at inaturalist.org, Donna Pomeroy.

An unusual sighting at Ilsley – a Yellow-throated Vireo. This photo is by Donna Pomeroy (CC-BY-NC) at inaturalist.org

We also saw the Red-eyed Vireo (Vireo olivaceus) again. Here’s a photo of this elusive bird by gifted local photographer, Bob Bonin.

The Red-eyed Vireo as photographed by gifted local photographer, Bob Bonin

Prairie Wildflowers – What a Sight!

Black-eyed Susan cover the northern edge of the eastern prairie as you walk in

In 2015, the eastern prairie was sown with native seed acquired through a grant from the U.S. Fish and Wildlife Service. It will take three or four summers of careful stewardship for these native, drought-resistant prairie plants to grow deep roots and reach full bloom. Although this is only their second summer, the native wildflowers are already carpeting the prairie with color.

Ben mowed the field once this year already and will mow once more in August.  This relatively high mowing reduces the seed production of annual and biennial plants (like burdock, thistle and Queen Anne’s Lace) and allows sunlight to reach the prairie plants while they are small and growing roots. Look  at just of few of the native wildflowers already coming up!

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And we still have autumn flowers to look forward to!

That Carpet of Prairie Wildflowers Brought All Kinds of Butterflies!

What a glorious sight! All over Ilsley’s eastern prairie, butterflies float and flutter in and out of the colorful tapestry of native plants – sipping the nectar and enjoying the sunlight. The birders sighted a relatively uncommon black butterfly called the Spicebush Swallowtail (Papilio troilus), though it took a lot of talk and research to identify it correctly! Luckily, the next day I saw the very similar Black Swallowtail (Papilio polyxenes) and was able to study the photos to spot the distinctions between the two. Here’s a swallowtail identification link I found very useful.  (Note that on this site,  the butterflies’ names are above rather than below the photos, which can be confusing at first.)

This is the male Spicebush Swallowtail. Notice that he only has white dots along the edge of his forewing, a faint line of pale blue dots above and a wash of blue (or blue scaling) on each hindwing.

A male Spicebush Swallowtail, a fairly rare butterfly to see in our area though they do frequent southern Michigan

The Black Swallowtail has a larger second (or postmedian) row of  yellowish spots above the row at the edge of its forewings and its more intense blue is contained in spots, rather than a wash of blue. It has a complete orange dot on each hindwing. This is the female, I believe, since her orange dots have a black center and the male’s are solid orange. Whew! Glad I had photos to help me!

The female Black Swallowtail, also at Ilsley, looks very similar to the male Spicebush.

My husband and I saw a third swallowtail flying high into the trees at Ilsley – the Eastern Tiger Swallowtail (Papilio glaucus). Here’s a photo from Bear Creek several years ago, just to remind you of its beauty, too. The female of this species can also be black!  But that’s for another blog…

Male Yellow Swallowtail butterfly closeup on bull thistle

One hot morning, four different Monarch butterflies (Danaus plexippus) drank from blossoms on the Eastern Prairie. They didn’t leave their wings open for long when they settled; they only used them to move from bloom to bloom. Perhaps closed wings kept them cooler, since less sunlight would fall on their closed wings than on open ones. I’ve learned that dragonflies tip their bodies vertically to stay cool on really hot days for that reason.

Some of the birders spotted a Red Admiral (Vanessa atalanta) one Wednesday morning when I was off looking at the Spicebush Swallowtail.  So here’s an older Bear Creek photo of one of those also.

Look for Red Admirals at Ilsley too!

On my last trip to the park, I saw a finger-nail-sized, triangular-shaped, white butterfly that I could not identify (help anyone?) and a tiny orange one that I believe is a Delaware Skipper (Anatrytone logan). To assess their size, consider that the little white one is under a Daisy Fleabane and the orange one is on a blade of grass! Interesting things come in small packages, eh?

A Spray of Tiny Grasshopper Nymphs,  a Nice Big Carolina Locust, and a Widow Skimmer Dragonfly as well

On the path that crosses the center of the Eastern Prairie, you’ll see – and hear! – hundreds of tiny grasshopper nymphs jumping off the path. They sound like rain falling on the taller grass stalks! Only about an half inch long now, they’re hard to identify. My guess is they’re Red-legged Grasshoppers (Melanoplus femurrubrum)The larger one in the slideshow below is definitely a Red-legged but it’s still a nymph, too. These guys molt 5-7 times before reaching adulthood. The brown Carolina Locust (Dissosteira carolina) is much bigger, about 2 inches.

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Of course no prairie is complete without dragonflies! Female, or possibly juvenile male, Widow Skimmers (Libellula luctuosa), with yellow abdominal stripes, hunted above the greenery last week. Adult males eventually develop a white wing band next to the dark one and their brown abdomens take on a bluish-white sheen as they mature.

A female or juvenile male Widow Skimmer dragonfly

Step by Step, Restoring Our Natural History

Eastern prairie at Charles Ilsley Park.

We all love local history, so it makes such perfect sense to restore our natural areas, too. At Charles Ilsley Park, we have an opportunity to see what our township looked like before European settlement, when Native American tribes lived on the rolling oak savannahs covered with tall grasses, wildflowers and widely spaced oaks. Even more importantly, natural areas restoration re-creates at least some of the rich diversity of plants, insects, birds and other wildlife that historically shared our green corner of the world. We’re on our way now to something very special here in Oakland Township.  Come have a look!

Footnote: My sources for information, besides Oakland Township's Stewardship Manager Ben VanderWeide, are as follows: butterfliesathome.com;Pollinators of Native Plants by Heather Holm; Stokes Nature Guides: A Guide to Bird Behavior Volumes 1-3; Allaboutbirds.org, the website of the Cornell Ornithology Lab at Cornell University; Wikipedia;  Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela; other sites as cited in the text.

Out and About in Oakland: Gallagher Creek Park – A Nature Oasis in a Busy Area

 

Viewing Platform GC
Viewing Platform at Gallagher Creek Park

I’ll admit that when I first visited Gallagher Creek Park, just east of Adams on Silverbell, I didn’t quite get it. It seemed like a rather ordinary, flat piece of land surrounded by a marsh. But luckily, I decided to look more closely and, as usual, the closer I look, the more nature shares with me. I’ve visited the park several times over the last month and each time I appreciate this little park more.

Blog post and photos by Cam Mannino
Blog post and photos
by Cam Mannino

The stream flowing quietly through the grass beneath the viewing platform creates a cool, moist retreat for an abundance of birds and the special native plants that love wet feet. The Old Fields bloom with an astonishing variety of native wildflowers. Insects float and buzz between the blossoms. In short, this little park hosts an amazing variety of wildlife and plants with its combination of marsh, wet meadows and open fields (click here to see a map).  Let me show you just a sampling.

 

 

Gallagher Creek Itself:  A Haven for Birds and Wildflowers

The recently constructed viewing platform (thanks to Eagle Scout Jonathan Walling!) near Gallagher Creek allows visitors to watch the slow ripples of cool water flowing through the grass and trees. This gentle flow creates a soothing retreat that shelters creatures and plants of all kinds. According to three surveys by the Michigan Department of Natural Resources, Gallagher Creek is home to “one of the few remaining self-sustaining Brook Trout (Salvelinus fontinalis) populations in southern Michigan.”

After the last survey, though, they reported that the “gravel riffles and natural pools” had been silted by nearby construction, reducing the population from 300 Brook Trout per mile in 1998 to just 50 per mile in 2010.  (See Ben’s blog on this from 2015.) Native plants growing more abundantly at the edges of the park can act as a buffer and filter such sediment. Gallagher Creek runs northeast and enters Paint Creek at the Cider Mill in Goodison. If we want Brook Trout in our township, one way is to protect a cold water stream like Gallagher Creek.

Wetland Wildlife and Plants: A Riparian Corridor and a Marsh

Streams like Gallagher Creek create what’s called a “riparian corridor,” an area where the land meets a stream or river that not only filters and purifies water, but also provides habitat for a wide diversity of plants and wildlife. Antonio Xeira, our birder friend, spotted a  most impressive bird  near the creek. Here’s his photo of a Great Horned Owl (Bubo virginianus) who spends its days high up among the leaves of a Willow (probably black willow, Salix nigra) north of the stream. It’s not easy to see so I appreciate Antonio’s shot!  See that open eye?

Great Horned Owl GC Antonio 8 2016
A Great Horned Owl, photographed by Antonio Xeira, spends the day in a large Willow tree to the north of Gallagher Creek.

A much smaller visitor among the willows is what I think was a Willow or Alder Flycatcher (Empidonax species) that perched in the shadows to preen. ( It could also be a Eastern Wood-Pewee (Contopus virens), but I’ll leave that for you to decide. They’re very similar.) It was a hot morning and this little bird was very busy dealing with the gnats or mites that can bother birds as much as they can bother us humans this time of year!

Willow Flycatcher
A Willow or Alder Flycatcher or an Eastern Wood-Pewee? You decide which one preened one hot morning in a Black Willow.

The shade near the stream also provided a stopover for a busy House Wren (Troglodytes aedon).

House Wren
A House Wren stops briefly in the willow’s shade on a hot morning.

This hard-working adult was foraging for her young in a nearby thicket of native Gray Dogwood (Cornus foemina), which provides lots of cover and a food source for all kinds of birds at Gallagher Creek.

Gray Dogwood GC
Stands of native Gray Dogwood provide shade, protection and a food source in several thickets around Gallagher Creek.

One morning, I approached the thicket to see if I could see the young wren.  Immediately, the adult starting her scolding call, chipping incessantly at me, probably both to ward me off and to tell her fledgling to stay back in the trees. I did finally manage to get a shot of the youngster who appeared to still be growing into its beak!

Fledgling wren GC
A fledgling House Wren hid deep inside the shade of Gray Dogwood as its mother scolded nearby.

Back in May, Antonio also noticed the burrow of a Crayfish (superfamilies Astacoidea and Parastacoidea),  a creature that loves clear, cold water streams. In case you haven’t seen a crayfish, I include a photo of one that I saw in Bear Creek.  If you enlarge the photo by clicking on it, you’ll see it’s a female carrying eggs under her tail! Crayfish are daylight creatures who retreat to burrows at night. I can’t say which species is in either photo. We do have one invasive crayfish in Michigan, the Rusty Crayfish (Orconectes rusticus), which fishermen used to use as bait and which have affected populations of our native crayfish.(Hover cursor for captions; click on photo to enlarge.)

Moisture-loving wildflowers in pink, purple, white and yellow bloom among the tall grasses and reeds  at the edge of the stream. Here are three native flowers that are easy to spot in the park – Blue Vervain (Verbena hastata), Canada Goldenrod (Solidago canadensis) and Joe Pye (Eutrochium maculatum).

The moist ground even a bit farther out from the stream suits many plants. Tall Sunflowers (Helianthus giganteus) and Water Hemlocks (Cicuta maculata), both natives,  prefer to sink their roots in moist earth.

Of course, dragonflies make their home wherever there is water. I saw a mating pair of Ruby Meadowhawks (Sympetrum rubicundulum) making a wheel of their bodies in the wet grass one morning.  The female is retrieving sperm from the male’s abdomen where he placed it earlier.  And to the right is what I think is another female Ruby Meadowhawk perching near the edge of the stream. Meadowhawk dragonflies (family Libellulidae)are plentiful this time of year and tough to identify,  but fascinating to watch as they hunt along the creek.

This thumbnail-sized Skipper butterfly (family Hesperiidae), I’m guessing a Delaware Skipper (Anatrytone logan), paused delicately on a moist leaf as I approached the stream.  Isn’t it elegant with its striped legs and antennae, large eyes and deep orange wings?

Delaware Skipper?
A thumbnail-sized member of the Skipper butterfly family, perhaps a Delaware Skipper.

Gallagher Creek Park also shares an emergent marsh with its neighbors to the east. This thriving area is slowly being cleared of the dreaded invasive, Phragmites (Phragmites australis), which is much too prevalent in the area. It’s an on-going project of Dr. Ben and the Parks and Recreation Commision. How wonderful to see natives like sedges (Carex species) and Joe Pye flourishing as that terrible invasive plant diminishes!

Emergent Marsh GC (1)
Native plants like Sedges and Joe Pye flourish in the park’s emergent marsh as the PRC works to eliminate invasive Phragmites.

Wildlife and Plants in the Old Fields

In the spring and early summer, our sharp-eyed birding friend, Antonio, found two nests on open ground at Gallagher Creek. I wonder if these eggs hatched since they seem so vulnerable. On the left is a Killdeer’s nest (Charadrius vociferus) from late April. The name, by the way, comes from their call, not from their effect on deer! And on the right is  a Spotted Sandpiper‘s nest (Actitis macularius) in early June.  Thanks, Antonio!

Now birds whisk in and out of the leafy trees at the perimeter of the Old Field next to the parking lot – many of them juveniles or adults working at feeding them. The “miaou” of a young Gray Catbird (Dumetella carolinensis) caught my attention at the edge of the field.  And later hearing a “cluck,” I look up to see an American Robin (Turdus migratorius), perhaps a molting female adult or a juvenile losing its spots, perched at the top of a snag.

A young Gray Catbird "miaous" from bushes near at the edge of the Old Field.
A young Gray Catbird “miaous” from bushes near the edge of the Old Field.
An immature Robin, still losing its breast spots, surveys Gallagher Creek.
A Robin, perhaps a molting female or a youngster still losing its breast spots, surveys Gallagher Creek.

Chipping Sparrow (Spizella passerina) and a male House Finch (Haemorhous mexicanus) hung out together in the bushes one hot morning.

Chipping sparrow and House Finch
A Chipping Sparrow and a House Finch share a bush one hot summer morning.

American Goldfinches (Spinus tristis) provide the soundtrack for Gallagher Creek right now. These cheery little birds are one of the few who sing as they fly! With every downward swoop of their undulating flight comes a short burst of song. Here’s a male that settled for a few moments in that Willow, again – a popular place to pause for many of the birds here.

Goldfinch in willow GC
A male goldfinch settles in the willow trees, resting from his singing flight in the golfinch mating season.

Goldfinches mate in August because their favorite food source is available – thistle seed.  So they’re quite happy, I imagine,  that non-native Field Thistle (Cirsium arvense) established a large, plumed patch on the western end  of the loop path.

Field thistle GC
Goldfinches mate in August when the Field thistles produce their favorite seed.

We appreciated being greeted by a relaxed Eastern Cottontail (Sylvilagus floridanus) when parking at Gallagher Creek one morning.  It seemed to be enjoying a bit of morning light after a long night of nibbling.

Rabbit GC
An Eastern Cottontail enjoys early morning sun after a long night of nibbling at Gallagher Creek.

As Reg and I set out on the path, we were greeted by a sight I’d never seen before.  A Downy Woodpecker (Picoides pubescens) hung onto one of the huge non-native stalks of Common Mullein (Verbascum thapsus) in the Old Field. Downies eat both insects and seeds, so I guess it was just varying its diet

Downy Woodpecer on Common Mullein
A Downy Woodpecker extracting seeds from a huge non-native plant, Common Mullein.

Closer to the water, this small bird fluffed its feathers as it settled in a bush. It looks a lot like the Willow Flycatcher but was a little bigger. So I’m guessing this is an Eastern Wood-Peewee (Contopus virens), but I can’t be sure!

Eastern Wood-PeeWee
I think from its size, this is an Eastern Wood-Pewee, but again, it could be a Flycatcher!

Among all the non-native wildflowers, like Queen Anne’s Lace and Spotted Knapweed in the Old Fields, many native wildflowers are increasingly making a home in the Old Fields here. In August of 2014,  Dr. Ben did what’s called a floristic survey of Gallagher Creek.  Of the 192 species he found on one day there, 131 were native!  Today they are more plentiful and healthy because Ben and his summer techs have eliminated many invasive shrubby plants and held repeated prescribed burns here. They plan to increase the native plant habitat in this park by planting native prairie species into the areas that were primarily non-native, invasive plants. The first phase of native plant installation will happen fall 2016, with a second round in 2017. Check out the sign on the west side of the driveway just as you enter the parking lot to learn more details.

In Bear Creek, Yarrow (Achillea millefolium) is an occasional plant, but  this summer, it grew in large swathes along the paths of the field near the parking lot at Gallagher Creek. Now, after the intense heat, it is turning brown and seeding for next spring’s crop.

Of course, native Common Milkweed (Asclepias syriaca) appears here and there, and is now producing its green, paisley-shaped pods. Along with more common Canada Goldenrod, another native called Grass-leaved Goldenrod (Euthamia graminifolia), flourishes as well. As I reminded readers last summer, Goldenrods do not cause “hay fever.” Their pollen is heavy and drops quickly to the ground. The sneeze-producing culprit that blooms at the same time is the wind-pollinated Ragweed (genus Ambrosia) which you can see if you click on the link.

Common Evening Primrose (Oenothera biennis) raises its graceful stems topped by yellow flowers among the grasses to the east of the park.  According to the Illinois Wildflower site (one of my faves), these flowers “remain open from evening to early morning, but will remain open longer on cloudy days. They have a mild lemony scent, and bloom from mid-summer to fall on mature plants. Long narrow seedpods develop, which split open from the top to release many tiny, irregular brown seeds. They are small enough to be dispersed by the wind, and can remain viable in the soil after 70 years.”  So if we keep caring for the land here, these night-blooming natives should be here for your grandchildren!

Common Evening Primrose blooms as the sun goes down until morning.
Common Evening Primrose open as the sun goes down  and don’t close until morning.

Out in the Old Fields, insects are busy in the late summer sun.  Carolina Locusts (Dissosteira carolina) spring into flight, their wings making a brown blur at your feet.

Grasshopper at GC
Carolina Locusts fly up as your tread the paths of Gallagher Creek.

Viceroy butterfly (Limenitis archippus) explored the possibilities of a Queen’s Anne’s Lace bloom one hot morning. Viceroys are distinguished from Monarchs by the bars on their hindwings.

Viceroy on Queen Anne's Lace
A Viceroy butterfly is distinguished from a Monarch by bars on its hindwings.

Common Ringlet (Coenonympha tullia) hovered in the grass below. Ringlets aren’t the most efficient flyers but they are prolific, occurring in Northern Europe, Asia and across North America but not in the southern hemisphere .

Common ringlet butterly
The Common Ringlet butterfly hovers close to the ground searching for nectar from small plants.

Clouded Sulphurs (Colias philodice) have a larger range of flight, feeding on taller plants like the Milkweeds and Coneflowers, but also on alfalfa or clover closer to the ground.

Clouded Sulphur on Spotted Knapweed
A Clouded Sulphur among the non-native Spotted Knapweed and Queen Anne’s Lace.

So, this modest little park is actually a rich oasis of native plants and wildlife along an important cold water stream  in the midst of the most developed area of our township.

Joe Pye GC
Joe Pye and Goldenrod complement each other near the creek where the reeds are growing in the background.

The Parks and Recreation Commission has plans to put a pavilion and perhaps even a play area in Gallagher Creek to make it even more inviting to its neighbors.  But right now, if you bring your binoculars and your curiosity to Gallagher Creek as the summer wanes, nature, as always, will reward you with lots of beauty to explore.

Footnote:  My sources for information are as follows: Ritland, D. B., & Brower, L. P. (1991); Stokes Nature Guides: A Guide to Bird Behavior Volumes 1-3, Allaboutbirds.org, the website of the Cornell Ornithology Lab at Cornell University; Wikipedia; http://www.butterfliesandmoths.org; Herbarium of the University of Michigan at michiganflora.net; various Michigan Field Guides by Stan Tekiela; Butterflies of Michigan Field Guide by Jaret C. Daniels; University of Wisconsin's Bug Lady at www4.uwm.edu/fieldstation/naturalhistory/bugoftheweek/ for insect info; http://www.migrationresearch.org/mbo/id/rbgr.html for migration info; invaluable wildflower identification from local expert, Maryann Whitman; experienced birder Ruth Glass, bird walk leader at Stoney Creek Metro Park for bird identification; Birds of North America Online; Audubon.org; Nature in Winter by Donald Stokes, Trees in My Forest by Bernd Heinrich, Winter World by Bernd Heinrich, Savannah River Ecology Lab (Univ of Georgia); Tortoise Trust website www.tortoisetrust.org;  An Orchard Invisible: A Natural History of Seeds by Jonathan Silvertown; The Ecology of Plants by Gurevitch, Scheiner and Fox; other sites as cited in the text.